Pleuroploca giganteus, with much confusion with its name, with Triplofusus papillosus now considered a nomen dubium, commonly known as the Florida horse conch although range stretches to Mexico. This is the second largest gastropod species, second only to the syrinx aruanus. The animal can retract the soft parts entirely into the shell and close it with the operculum.

The soft parts are bright orange in color. The size of the shell can reach over 600mm! The shell color is bright orange in very young individuals. The shell often becomes greyish white to salmon-orange when adult, with a light tan or dark brown periostracum. This species dwells on sand, weed and mud flats from the low intertidal to shallow subtidal zones, in 20 foot (6 m) deep water.

Triplofusus papillosus is a carnivorous predatory species, and feeds on other large marine gastropods, including the tulip shell (Fasciolaria tulipa), the lightning whelk (Sinistrofulgur perversum), and the queen conch (Lobatus gigas) as well as some Murex species. It may also present cannibalistic behaviour, feeding on smaller conspecific individuals. It has been observed (in an aquarium setting) to eat small hermit crabs of the species Clibanarius vittatus.

The U.S. state of Florida declared it the state seashell in 1969. The shell is popular with shell collectors partly because of its great size.

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